Choice B is the best answer. The second sentence of the last paragraph says that Alexander Volkov and his colleagues "first demonstrated that it is indeed electricity that causes the Venus flytrap to close." In this context, the word "demonstrated" most nearly means established.
Choices A, C, and D are incorrect because in the context of scientists showing what causes the Venus flytrap to close, the word "demonstrated" most nearly means established, not protested (choice A), performed (choice C), or argued (choice D).