Choice C is the best answer. When discussing problems with bee colo-nies, the authors use phrases like "we suspect" (line 19"We... bee") and "we postulate" (line 21"deficient...problem") to show they are hypothesizing reasons for bee colonies' suscep-tibility to mite infestations. The use of "can," "may," and "could" creates a tentative tone and provides further evidence that the authors believe, but are not certain, that their hypothesis is correct.
Choices A, B, and D are incorrect because the authors' use of "can," "may," and "could" does not create an optimistic, dubious, or critical tone.